In the case of two subpopulations, . Equation (8) then simplifies to
This system and its solution were derived by Nath and Griffiths (1993). Setting M = 0 and solving the system for , and gives the expected pairwise coalescence times of motif 2 (). Considering M > 0 and solving the system gives the expected pairwise coalescence times of motif 3 ().
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