We examined substitution rates synonymous (Ks) and non-synonymous (Ka) using Model Averaging in the KaKs_Cal-culator program (Institute of Genomics, Chinese Academy of Sciences, Beijing, China) [64]. Eighty common protein-coding genes shared by the I. pinfanensis and H. triflora were aligned separately using Geneious software v5.6.4 (Biomatters Ltd., Auckland, New Zealand) [59].
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