Animal Subjects.

Lister-Hooded rats (n = 8) were 2 to 3 mo old at the start of the experiments. Animals were purchased from Charles River and group housed until surgery; after surgery, animals were housed in single cages. During the behavioral task, they were food deprived to 85 to 90% of the free-feeding weight against a growth curve, with free access to water, on a 12:12-h light-dark cycle with training in the light phase. Care of the animals complied with the UK Animals (Scientific Procedures) Act conducted under a Project License (PPL P7AA53C3F). Two animals were excluded from the recordings due to insufficient quality of the field of view (FOV; low number of detected cells). One further animal (H0487) could only be recorded in sessions 18 and 19 due to technical issues and was excluded from the analysis on prospective coding because we could not compare the three possible destinations.