Cells were incubated under dark, anaerobic conditions as wells as the experiment of organic acid production. The accumulated H2 gas in the headspace of a GC-vial was measured with a gas chromatograph (GC-2010 Plus AT, Shimadzu, Kyoto, Japan) as described previously24 and in accordance with the manufacturer’s instructions. N2 was used as the carrier gas with a flow rate of 10 mL/min.
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