A composite interval mapping analysis was carried out using R/qtl software for QTL mapping [52]. The threshold of the QTL value was determined at a significance level of P < 0.05 after 1,000 permutations. To investigate the genetic overlap between each trait, the 2-LOD support intervals of QTL were compared; QTLs with overlapping support intervals were regarded as common QTLs for the compared traits.
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