Further information and requests for resources and reagents should be directed to and will be fulfilled by the Lead Contact, Zihe Rao (raozh@tsinghua.edu.cn).

This study did not generate new unique reagents.

The cryo-EM density maps and the structures were deposited into the Electron Microscopy Data Bank (EMDB) and Protein Data Bank (PDB) with the accession numbers 31138 and 7EGQ for dCap(0)-RTC, and 31146 and 7EIZ for mCap(0)-RTC.

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