To quantify the phase shift in the output PC spike train relative to the frequency of sinusoidal modulation of GC input spike train, the average PC firing rate over the whole population was fitted by a sinusoidal function Asin(2πft + ϕ) + C, where C is the offset of firing rate, and the modulation phase shift at frequency f is thus fully specified by the phase shift ϕ of the sinusoidal component, since the initial phase of input stimulus was set as 0 in all simulations.

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