Resources, reagents and further information requirement should be forwarded to and will be responded by the Lead Contact, David I Stuart (dave@strubi.ox.ac.uk).

Reagents generated in this study are available from the Lead Contact with a completed Materials Transfer Agreement.

The coordinates and structure factors of the SARS-CoV-2 RBD-N501/Fab269 and SARS-CoV-2 RBD-501Y/Fab269 crystallographic complexes are available from the PDB with accession codes PDB: 7NEH,7NEG, respectively. Mabscape is available from https://github.com/helenginn/mabscape, https://snapcraft.io/mabscape. The data that support the findings of this study are available from the corresponding authors on request.

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