The identified lncRNAs were further classified as intergenic, intronic, sense, and antisense lncRNAs based on the spatial relationships of their transcripts loci with PCGs using Cuffcompare tool [50]. The transcripts length and exon number for protein-coding transcripts and lncRNA transcripts were assessed [52]. The proportion of different kinds of lncRNAs and protein-coding transcripts was calculated.

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