Domains were identified based on conservation calls above and examined using the UCSC browser to compare the arrangement of genes. TAD boundaries were taken from Dixon, 2012 [36] and were annotated to genes if they were between it and the next gene or were within the gene body. Additionally, to confirm that UBA1 TSSs were within the same TAD, we used a larger set of TADs in the 3D genome browser [69].

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