Platanus Genome Assembler (v1.2.4)13 was used to assemble all the high-quality reads into scaffolds with parameters “-c 15 -k 60 -t 50 -m 300”. GapCloser ( was adopted with parameters “-p 25 -l 150” to fill gaps in the assembled scaffolds using PE reads. Read mapping, BUSCO, CEGMA, and BAC alignment were used to evaluate the quality of the assembled genome.

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