Lipids 1,2-dioleoyl-sn-glycero-3-phosphocholine (DOPC), brain l-α-PS, and brain PI(4,5)P2 were purchased from Avanti Polar Lipids (Alabaster, AL, USA). ATTO647N-DPPE and ATTO647N–SM (ATTO647N-SM) were purchased from ATTO-TEC GmbH (Siegen-Weidenau, Siegen, Germany). Abberior STAR RED (KK114)–1,2-dihexadecanoyl-sn-glycero-3-phosphoethanolamine (KK114-DPPE), cholesterol–PEG(1 k)-Abberior STAR RED (KK114) [Chol-PEG(1 k)-KK114], and ATTO647N-PI(4,5)P2 [ATTO647N-PI(4,5)P2] were all purchased from Abberior GmbH (Göttingen, Germany). Chemical structures of all the lipid analogs used in this study are given in fig. S1.

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