For the grip-strength test, each rat was lifted by its tail and its forelimbs (or all limbs) were allowed to grasp metal mesh fixed to a force-electricity transducer (Brain Science Idea Co. Ltd.; BS-TM-RM). The rat was gently pulled upward while it grasped the mesh with its forelimbs (or all limbs). The maximal force reached immediately before the rat released the mesh was taken as the grip strength.

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