Systematic literature wide opened search was performed in electronic databases, also including the grey literature ( Figure 1 ). General controlled vocabulary (MeSH Terms) and keywords were used and the searches had no language, year, or publication type restriction. The main terms included “microbiota”, “microbiome”, “human microbiota”, “low abundant”, “minority species”, “keystone”. The search strategy and the results retrieved in each electronic database are shown in Appendix 1 . Duplicated references were removed by the reference manager EndNoteWeb (Clarivate Analytics, Mumbai) and then manually.

Flow diagram for study selection according to PRISMA guidelines. *Some studies sampled multiple sites in one study.

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